1 Biological Sciences, Northern Illinois University, 1425 W Lincoln Hwy, DeKalb 60115-2861, IL, USA
2 Ecology, Evolution and Organismal Biology, Iowa State University, 251 Bessey Hall, Ames 50011-1020, IA, USA
3 Instituto de EcologÃa AC, Centro Regional del BajÃo, Biodiversidad y Sistemática, Av. Lázaro Cárdenas 253, Pátzcuaro 61600, Michoacán, Mexico