1 Department of Ophthalmology and Visual Sciences, John A Moran Eye Center, University of Utah, 65 Mario Capecchi Drive, Salt Lake City 84132, UT, USA
2 Interdepartmental Program in Neuroscience, University of Utah, 20 North 1900 East, Salt Lake City 84132, UT, USA
3 Department of Neurobiology and Anatomy, University of Utah, 20 North 1900 East, Salt Lake City 84132, UT, USA