1 Department of Biology, University of Copenhagen, Copenhagen, DK-2100, Denmark
2 Department of Biomedical Sciences, University of Copenhagen, Copenhagen, DK-2200, Denmark
3 Department of Systems Biology, Center for Biological Sequence Analysis, Technical University of Denmark, Kongens Lyngby, DK-2800, Denmark
4 Department of Veterinary Clinical and Animal Sciences, University of Copenhagen, Frederiksberg, DK-1870, Denmark
5 BGI-Shenzhen, Shenzhen 518083, China
6 Princess Al Jawhara Center of Excellence in the Research of Hereditary Disorders, King Abdulaziz University, Jeddah 21589, Saudi Arabia
7 Macau University of Science and Technology, Avenida Wai Long, Taipa, Macau 999078, China