1 Centre for Sustainable Tropical Fisheries and Aquaculture, College of Marine and Environmental Sciences, Building 34, James Cook University, Townsville 4811, Queensland, Australia
2 AIMS@JCU, Australian Institute of Marine Science, College of Marine and Environmental Sciences, Building 17, James Cook University, Townsville 4811, Queensland, Australia
3 Australian Institute of Marine Science, PMB No 3, Townsville MC, Townsville 4810, Queensland, Australia