1 Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China
2 University of Chinese Academy of Sciences, Beijing 100049, China
3 Department of Botany, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba 305-0005, Ibaraki, Japan
4 Jodrell Laboratory, Royal Botanic Gardens Kew, Richmond, Surrey TW9 3AB, UK
5 Yunnan Academy of Forestry/Yunnan Laboratory for Conservation of Rare, Endangered and Endemic Forest Plants, State Forestry Administration, Kunming 650204, Yunnan, China